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Ensure the software on all your desktops and servers with the latest service packs and updates.
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Implement a plan for regular updates, and automate it as much as possible.
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Deploy antivirus and anti-spyware on all your computers. Enable host-based firewalls.
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Physically secure your servers. A locked room is best; heavy cables are better than nothing.
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Plan for rolling out roaming profiles and folder redirection (with offline folders) so that client computers can be rebuilt with ease.
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Upgrade your wireless networking to WPA.
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Change all your passwords to pass phrases.
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Implement a backup plan; don't forget to regularly test the media.
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Write an Internet acceptable use policy and have all your employees sign it.